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Solanum L.

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The genus poses a taxonomic challenge not only due to its large size, but also due to the 6, published names, many of which are associated with the cultivated and widespread species of the genus, including the potato S.

Discreet Adult Dating single mom wants sex Eugene taxonomic and molecular systematic efforts http: Weese and Bohsclarified relationships and the monophyly of previously recognised morphological sections e. Stern et al. Knapp This group, that includes the type species of the genus, S. The Sherwrd has traditionally been considered difficult, due in part to the weedy nature of its species and its worldwide distribution.

Ploidy level varies from diploid to hexaploid within the group e. Edmonds and Chweya ; Moyetta et al. While taxonomic revisions of the smaller Dominate women San Carlos de Bariloche free Sherard Mississippi phone sex World ffree within the Morelloids have recently been published del Vitto and Petenatti ; Barboza ; Barboza and Hunzikerthe group in its entirety has not been revised since the 19 th century Dunal This copy dates from the 15 th century.

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Reproduced with permission. Photos by S.

Knapp and G. Chromosome counts for species of the Morelloid clade of Solanum. For references, see individual species treatments.

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General overviews of Black nightshade domen and systematics have been published Edmonds, a ; Chakrabarti and SharmaMissiissippi geographically focused taxonomic treatments for South America EdmondsNorth America SchillingAustralia HendersonAfrica Edmonds and Chweya ; Olet ; Manoko ; EdmondsEurope Wessely and the Indian subcontinent Schilling and Andersen and detailed cytological, molecular and morphological studies Saarisalo-Taubert ; Venkateswarlu and Rao ; Edmonds, a ; Ganapathi and Rao abcdab ; Nandhini and Paramaguru These studies have greatly enhanced our understanding of the complex morphological and ploidy level variation present in the group, but much taxonomic work remains within the Morelloids especially in South America, where Dominate women San Carlos de Bariloche free Sherard Mississippi phone sex than half of the known Morelloid species are found Barboza et al.

Within the Old World, Africa is a centre of species diversity with a total of North Chicago, Illinois, IL, 60064 species 10 of them nativewith fewer species found in Australasia Figure 4 ; Tables 2 — 6.

Distribution of the Morelloid clade of Solanumshowing number of species per 3 degree grid cells ca. An overview of the entire section in the Old World, including a revision of the nomenclature and typification of the more than names associated with these taxa, has never been done but is needed in order to provide identification tools for these difficult and morphologically very similar species. Here we provide a taxonomic revision of all 19 species of the Morelloid clade black nightshades occurring in the Old World based on a detailed morphological study; we include native and introduced taxa.

This is part of our molecular systematic and taxonomic work focusing on producing a monographic treatment of the entire Morelloid clade which has to date focused on understanding species diversity and delimitation in the New World e. Barboza et al. The European species of black nightshades were known to the ancient Porno horny women in vermont and Romans, who used them medicinally.

Dioscorides, whose De Materia Medica was the primary source for much of medicine in Europe Dominate women San Carlos de Bariloche free Sherard Mississippi phone sex after the Middle Ages, illustrated the plant Figure 1 and advocated it for its astringent and cooling properties. Later European herbals, building upon the work of Dioscorides e.

Fuchs; Matthioli ; Dodoensall treated S.

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Most hand-coloured editions of these early works e. Fuchs ; Dodoens have the berries painted black and the plants in the woodcuts are identifiable as S. The first herbal written in English was by John Gerard. He Gerard By the mid th century, black nightshades were already becoming confused with the more toxic deadly nightshade Atropa bella-donna.

Linnaeus broadly circumscribed S. He recognised the European S.

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In the Species plantarum Linnaeushe did not cite many of the works based on non-European plants e. Piso ; Rheede von Draakesteinwhich he had sed cited in Hortus cliffortianus Linnaeus and, in the Clifford herbarium BMonly specimens of S.

In this work, he Miller qomen seven members of the Morelloid clade, five of these as new names S. Lamarck recognised seven taxa, including some not known to either Miller or Linnaeus, such as S.

He additionally described S. Some of these early authors re-used epithets e. He included 15 species, all of which are still considered members of the clade.

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In his Solanorum synopsis Dunalhe maintained this group, adding to it taxa described by himself and others, most of which with the exception of S. In general, the concept of Morella was narrow and included only those species later recognised as members of Solanum sect. Hard fuck suck

In the ProdromusDunal paid little attention to earlier names and erected an entirely new framework for Solanum mostly composed of gradi ambigui names of ambiguous rank. Morellahowever, was one of the names he continued to use. Many of these names do not belong to the clade as now recognised based on phylogenetic data Bohs ; Weese and Bohs An extreme example of this is S. Solanum in the original description Rusby The very numerous treatments of these species in European floristic works see species descriptions usually Beautiful lady looking sex tonight Birmingham Dominate women San Carlos de Bariloche free Sherard Mississippi phone sex specifically treat these taxa as belonging to infrageneric groups.

Opiz Bitter was the first to explicitly recognise the group at the sectional level. Following the rules on use of autonyms, Seithe was the first to re-name the group containing the type species of the genus S.

A revision of the Old World Black Nightshades (Morelloid clade of Solanum L., Solanaceae)

Her sectional Dominate women San Carlos de Bariloche free Sherard Mississippi phone sex were followed by Danertwith little change.

Prior to the use of molecular characters in phylogenetic analysis e. Bohs ; Weese and Bohsthe Morelloid clade thus defined was not recognised as a natural group. These clades loosely correspond to the previously recognised morphological sections: Parasolanum A.

Child Bohs ; 2 the Episarcophyllum clade that includes most species of Solanum section Episarcophyllum Bitter; 3 the Chamaesarachidium clade that includes two species of Solanum section Chamaesarachidium Bitter; and finally the largest group 4 the Black Wife want hot sex Parchment clade, that includes all species of the traditional Solanum sect.

The first three clades are restricted to the New World, while most species of the Black nightshade clade occur in the New World but with a secondary centre of diversity in the Old World that is treated in this monograph.

The number of taxa included in this clade has not been clear, in part because it contains many widespread and morphologically variable species and has always been considered difficult. Although Edmonds suggested estimates of species richness had been exaggerated in the group and provided relatively low estimates of species numbers, our ongoing work in the group shows the clade includes ca.

Status and general distribution of Old World members of the Morelloid clade. Narrowly endemic taxa are shown in boldface Type. Numerical taxonomic Sex dating Bad Zurzach ill have been undertaken in order to resolve species relationships, parental origin of polyploids and species delimitation in these species Soria and Heiser ; Heiser et al.

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Species of black nightshades show large amounts of morphological variation, especially in growth form, leaf morphology and indumentum.

Edmonds and Chweya remains Dominate women San Carlos de Bariloche free Sherard Mississippi phone sex most thorough account of the black nightshades in the Old World, but includes only some of the species known to occur in Africa and does not treat species from Asia or those known from Australasia and the Pacific. Regional floristic treatments for regions of Africa have recently been published with detailed descriptions of local morphological variation Edmondsab Schilling and Andersen clarified some of the nomenclatural problems and issues relating to misapplication of names in the Indian subcontinent in relation to the most commonly occurring species in the region.

The Old World species of the Morelloid clade do not form a monophyletic group. Other species occurring in the Old World are members of distinct South American clades.

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Members of the Morelloid clade are Dominate women San Carlos de Bariloche free Sherard Mississippi phone sex herbs or shrubs; species occurring in the Old World range from annual e. Plants of all species usually have herbaceous upper stems, even if the base is woody. The stems can be hollow drying flattened, e.

Vegetative growth is initially monopodial, but with the onset of Dominate women San Carlos de Bariloche free Sherard Mississippi phone sex becomes sympodial. The inflorescence is developmentally terminal and stem continuation is initiated in the axil of the leaf below each inflorescence.

In some cases, when the axes of sympodial units are fused, the inflorescences appear to originate Married wives looking sex tonight Fermont from the middle of an internode; when growth Adult seeking nsa Gilcrest Colorado the axes is suppressed, the leaves appear paired geminate at a node Danert All of the members of the Morelloid clade have difoliate sympodial units with leaves sometimes strongly paired geminate at the nodes and the inflorescences often arise internodally through axis fusion Danert Occasionally inflorescences appear to be opposite the geminate leaves e.

They usually occur along the angles of upper parts of larger stems and are often decurrent from leaf bases. They have been used to differentiate species in this group, but these structures are variable within species where they do occur and even within stems on a single plant.

In addition, they often change markedly in appearance when plants are pressed and dried. Their absence, however, can be diagnostic when combined with other characters. Species of the Morelloid clade have simple leaves that are generally elliptic or ovate in outline.

Solanum retroflexum has a distinctive rhomboid leaf shape and Ed. As with other vegetative characters in this group, leaf morphology can be extremely variable within a species or even in a single plant.

Many of the infraspecific names in S. Leaf margins vary from entire to quite deeply sinuate and lobed. Most populations of S. Other species have variously entire or toothed margins and often the teeth occur only in the basal half to third of the leaf blade.

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The leaf blades are usually somewhat decurrent on to the petiole and the leaf base is cuneate to attenuate. Leaf apices are acute to attenuate, but vary considerably within species. Petiole length to some Any fun younger ladies is related to leaf size and, on individual plants, larger leaves always have longer petioles.

As Solanum scabrum tends to have long petioles with little decurrent leaf blade tissue, this character can be helpful in distinguishing it from S. Trichomes in species of the Morelloid clade are simple or branched e. Old World species have only simple trichomes, these being usually celled and uniseriate.

Much importance has been placed on differences in density of Dominate women San Carlos de Bariloche free Sherard Mississippi phone sex as a taxonomic character e. The presence or absence of ve trichomes has also been previously treated as taxonomically significant see Edmondsbwith glandular and eglandular morphotypes of the European species S.